I study the evolution of complex systems. Current student research includes investigations on the formation of ecological communities, developmental evolution, and the evolution of multicellularity. Our studies are largely experimental, observing ‘evolution-in-action’ using microbial systems. This work is highly collaborative and our extended research group includes EEB faculty Ford Denison, Mark Borrello & Tony Dean, and faculty from the Minnesota Center for the Philosophy of Science and Dept. of Plant Biology.